Updated Oct 28, 2026
DLA Class I Haplotype Frequencies
| DLA1 # | STR types | Havanese (n=1184) |
|---|---|---|
| 1003 | 387 375 277 186 | 0.0384 |
| 1006 | 387 375 293 180 | 0.0562 |
| 1009 | 382 377 277 184 | 0.0004 |
| 1011 | 376 365 281 180 | 0.0008 |
| 1012 | 388 369 289 188 | 0.0110 |
| 1014 | 375 373 287 178 | 0.0186 |
| 1016 | 382 371 277 178 | 0.1921 |
| 1017 | 386 373 289 178 | 0.0008 |
| 1018 | 375 373 287 186 | 0.0068 |
| 1019 | 380 373 287 185 | 0.0008 |
| 1029 | 380 365 281 182 | 0.0355 |
| 1030 | 380 373 293 178 | 0.0025 |
| 1033 | 382 379 277 181 | 0.0013 |
| 1035 | 386 373 277 184 | 0.0038 |
| 1040 | 380 371 277 186 | 0.0160 |
| 1052 | 380 372 289 184 | 0.0089 |
| 1054 | 382 379 277 184 | 0.1225 |
| 1062 | 382 371 277 183 | 0.0004 |
| 1068 | 380 373 287 181 | 0.0127 |
| 1071 | 380 373 277 178 | 0.0004 |
| 1084 | 376 373 277 184 | 0.0013 |
| 1087 | 380 371 277 178 | 0.0034 |
| 1092 | 376 379 277 181 | 0.0963 |
| 1093 | 386 379 277 180 | 0.0135 |
| 1105 | 382 379 277 178 | 0.0008 |
| 1111 | 387 378 287 182 | 0.0025 |
| 1114 | 380 373 287 183 | 0.0169 |
| 1115 | 386 371 277 182 | 0.0925 |
| 1116 | 380 365 289 186 | 0.1085 |
| 1117 | 376 373 277 180 | 0.0473 |
| 1118 | 376 377 277 180 | 0.0008 |
| 1120 | 376 386 289 176 | 0.0008 |
| 1121 | 380 371 277 183 | 0.0046 |
| 1123 | 386 379 277 184 | 0.0013 |
| 1124 | 388 373 289 178 | 0.0055 |
| 1125 | 393 383 277 185 | 0.0220 |
| 1128 | 384 376 287 182 | 0.0008 |
| 1129 | 382 371 277 181 | 0.0008 |
| 1132 | 376 379 277 184 | 0.0013 |
| 1133 | 378 365 287 172 | 0.0169 |
| 1134 | 384 365 291 178 | 0.0008 |
| 1136 | 382 371 277 182 | 0.0004 |
| 1140 | 376 379 301 180 | 0.0038 |
| 1142 | 376 379 277 180 | 0.0046 |
| 1147 | 391 375 293 180 | 0.0025 |
| 1148 | 376 375 277 180 | 0.0021 |
| 1154 | 376 365 281 183 | 0.0097 |
| 1155 | 388 369 287 184 | 0.0004 |
| 1173 | 392 371 277 186 | 0.0038 |
| 1180 | 386 371 277 181 | 0.0004 |
| 1260 | 387 379 277 184 | 0.0004 |
| 1262 | 382 377 289 180 | 0.0008 |
| 1266 | 378 365 287 181 | 0.0008 |
| 1290 | 380 369 277 181 | 0.0008 |
| 1300 | 388 369 260 184 | 0.0004 |
DLA Class II Haplotype Frequencies
| DLA2 # | STR types | Havanese (n=1184) |
|---|---|---|
| 2001 | 343 324 284 | 0.0427 |
| 2003 | 343 324 282 | 0.2141 |
| 2005 | 339 322 280 | 0.0025 |
| 2006 | 339 325 280 | 0.0013 |
| 2007 | 351 327 280 | 0.0587 |
| 2008 | 339 327 276 | 0.0004 |
| 2011 | 345 322 284 | 0.0004 |
| 2012 | 345 322 280 | 0.0025 |
| 2014 | 339 322 284 | 0.0034 |
| 2016 | 339 323 284 | 0.0266 |
| 2017 | 343 322 280 | 0.0097 |
| 2018 | 339 324 284 | 0.0249 |
| 2021 | 339 324 268 | 0.0046 |
| 2022 | 339 327 282 | 0.1153 |
| 2023 | 341 323 282 | 0.0025 |
| 2024 | 343 323 280 | 0.0034 |
| 2028 | 345 327 288 | 0.0008 |
| 2032 | 339 323 280 | 0.0182 |
| 2033 | 339 323 282 | 0.0013 |
| 2035 | 341 323 280 | 0.0021 |
| 2037 | 341 327 280 | 0.0186 |
| 2040 | 345 327 280 | 0.0038 |
| 2053 | 343 324 280 | 0.0287 |
| 2066 | 339 324 280 | 0.1639 |
| 2068 | 339 327 284 | 0.0004 |
| 2070 | 347 324 282 | 0.1081 |
| 2071 | 339 322 286 | 0.0046 |
| 2072 | 339 325 282 | 0.0169 |
| 2073 | 339 327 286 | 0.0084 |
| 2074 | 341 324 284 | 0.0473 |
| 2075 | 341 327 282 | 0.0363 |
| 2076 | 345 322 282 | 0.0055 |
| 2077 | 347 325 286 | 0.0169 |
| 2079 | 343 323 278 | 0.0008 |
| 2082 | 339 325 268 | 0.0038 |
| 2087 | 347 324 280 | 0.0004 |
Allele Frequencies
| # | Locus Name | Allele | Havanese (n=1200) |
|---|---|---|---|
| 1 | AHT121 | 88 | 0.0004 |
| 92 | 0.1067 | ||
| 94 | 0.0775 | ||
| 96 | 0.0021 | ||
| 98 | 0.0096 | ||
| 100 | 0.0471 | ||
| 102 | 0.1058 | ||
| 104 | 0.5296 | ||
| 106 | 0.0350 | ||
| 108 | 0.0821 | ||
| 110 | 0.0013 | ||
| 112 | 0.0029 | ||
| 2 | AHT137 | 131 | 0.2342 |
| 133 | 0.0621 | ||
| 137 | 0.0817 | ||
| 139 | 0.0400 | ||
| 141 | 0.1783 | ||
| 143 | 0.1121 | ||
| 145 | 0.0079 | ||
| 147 | 0.1988 | ||
| 149 | 0.0650 | ||
| 151 | 0.0192 | ||
| 155 | 0.0008 | ||
| 3 | AHTH130 | 117 | 0.0358 |
| 119 | 0.2671 | ||
| 121 | 0.0500 | ||
| 123 | 0.0317 | ||
| 125 | 0.3667 | ||
| 127 | 0.1850 | ||
| 129 | 0.0408 | ||
| 131 | 0.0200 | ||
| 133 | 0.0029 | ||
| 4 | AHTh171-A | 217 | 0.0017 |
| 219 | 0.1617 | ||
| 221 | 0.0013 | ||
| 225 | 0.2550 | ||
| 227 | 0.0304 | ||
| 229 | 0.1863 | ||
| 231 | 0.2771 | ||
| 233 | 0.0204 | ||
| 235 | 0.0383 | ||
| 237 | 0.0279 | ||
| 5 | AHTh260 | 238 | 0.1233 |
| 242 | 0.0133 | ||
| 244 | 0.2333 | ||
| 246 | 0.2888 | ||
| 248 | 0.2038 | ||
| 250 | 0.1054 | ||
| 252 | 0.0025 | ||
| 254 | 0.0279 | ||
| 258 | 0.0017 | ||
| 6 | AHTk211 | 87 | 0.4746 |
| 89 | 0.2258 | ||
| 91 | 0.1454 | ||
| 95 | 0.1429 | ||
| 97 | 0.0113 | ||
| 7 | AHTk253 | 284 | 0.0117 |
| 286 | 0.0696 | ||
| 288 | 0.6442 | ||
| 290 | 0.0488 | ||
| 292 | 0.2258 | ||
| 8 | C22.279 | 116 | 0.1029 |
| 118 | 0.2121 | ||
| 120 | 0.0050 | ||
| 122 | 0.0171 | ||
| 124 | 0.2308 | ||
| 126 | 0.1329 | ||
| 128 | 0.0008 | ||
| 130 | 0.2983 | ||
| 9 | FH2001 | 124 | 0.0404 |
| 128 | 0.0092 | ||
| 132 | 0.4117 | ||
| 136 | 0.0113 | ||
| 140 | 0.0113 | ||
| 144 | 0.4533 | ||
| 148 | 0.0513 | ||
| 152 | 0.0100 | ||
| 158 | 0.0017 | ||
| 10 | FH2054 | 148 | 0.0158 |
| 152 | 0.0842 | ||
| 156 | 0.2513 | ||
| 160 | 0.1979 | ||
| 164 | 0.0879 | ||
| 168 | 0.1458 | ||
| 172 | 0.1113 | ||
| 176 | 0.1021 | ||
| 180 | 0.0033 | ||
| 184 | 0.0004 | ||
| 11 | FH2848 | 230 | 0.0138 |
| 232 | 0.3042 | ||
| 234 | 0.0483 | ||
| 236 | 0.0817 | ||
| 238 | 0.0463 | ||
| 240 | 0.2333 | ||
| 242 | 0.2625 | ||
| 244 | 0.0058 | ||
| 246 | 0.0042 | ||
| 12 | INRA21 | 91 | 0.1375 |
| 95 | 0.0992 | ||
| 97 | 0.3758 | ||
| 99 | 0.0500 | ||
| 101 | 0.1775 | ||
| 103 | 0.1492 | ||
| 105 | 0.0108 | ||
| 13 | INU005 | 106 | 0.0008 |
| 110 | 0.0308 | ||
| 122 | 0.0133 | ||
| 124 | 0.5871 | ||
| 126 | 0.2175 | ||
| 128 | 0.0088 | ||
| 132 | 0.0275 | ||
| 134 | 0.0046 | ||
| 138 | 0.1096 | ||
| 14 | INU030 | 144 | 0.6321 |
| 146 | 0.0392 | ||
| 148 | 0.0238 | ||
| 150 | 0.1271 | ||
| 152 | 0.1633 | ||
| 154 | 0.0133 | ||
| 156 | 0.0013 | ||
| 15 | INU055 | 210 | 0.3742 |
| 212 | 0.2058 | ||
| 214 | 0.1004 | ||
| 216 | 0.1300 | ||
| 218 | 0.1842 | ||
| 222 | 0.0054 | ||
| 16 | LEI004 | 83 | 0.0004 |
| 85 | 0.6408 | ||
| 93 | 0.0004 | ||
| 95 | 0.2258 | ||
| 97 | 0.0146 | ||
| 101 | 0.0017 | ||
| 105 | 0.0004 | ||
| 107 | 0.1088 | ||
| 109 | 0.0071 | ||
| 17 | REN105L03 | 227 | 0.0221 |
| 229 | 0.0183 | ||
| 231 | 0.2096 | ||
| 233 | 0.0925 | ||
| 235 | 0.0042 | ||
| 237 | 0.1900 | ||
| 239 | 0.0113 | ||
| 241 | 0.4463 | ||
| 245 | 0.0058 | ||
| 18 | REN162C04 | 200 | 0.0346 |
| 202 | 0.2196 | ||
| 204 | 0.0492 | ||
| 206 | 0.3971 | ||
| 208 | 0.1288 | ||
| 210 | 0.1708 | ||
| 19 | REN169D01 | 202 | 0.0692 |
| 204 | 0.0008 | ||
| 210 | 0.1679 | ||
| 212 | 0.0525 | ||
| 214 | 0.0267 | ||
| 216 | 0.4192 | ||
| 218 | 0.0704 | ||
| 220 | 0.1933 | ||
| 20 | REN169O18 | 160 | 0.0708 |
| 162 | 0.2042 | ||
| 164 | 0.0438 | ||
| 166 | 0.2317 | ||
| 168 | 0.1850 | ||
| 170 | 0.2342 | ||
| 172 | 0.0304 | ||
| 21 | REN247M23 | 266 | 0.0004 |
| 268 | 0.4113 | ||
| 270 | 0.1029 | ||
| 272 | 0.1663 | ||
| 274 | 0.0188 | ||
| 276 | 0.2708 | ||
| 278 | 0.0296 | ||
| 22 | REN54P11 | 222 | 0.1688 |
| 226 | 0.2663 | ||
| 228 | 0.1117 | ||
| 232 | 0.0900 | ||
| 234 | 0.0596 | ||
| 236 | 0.0413 | ||
| 238 | 0.2625 | ||
| 23 | REN64E19 | 139 | 0.1413 |
| 143 | 0.0133 | ||
| 145 | 0.0983 | ||
| 147 | 0.3108 | ||
| 149 | 0.0150 | ||
| 151 | 0.0038 | ||
| 153 | 0.3996 | ||
| 155 | 0.0175 | ||
| 157 | 0.0004 | ||
| 24 | VGL0760 | 12 | 0.0317 |
| 13 | 0.0608 | ||
| 14 | 0.1029 | ||
| 15 | 0.0096 | ||
| 18.2 | 0.0971 | ||
| 19.2 | 0.0025 | ||
| 20.2 | 0.0171 | ||
| 21.2 | 0.2333 | ||
| 22.2 | 0.1704 | ||
| 23.2 | 0.1608 | ||
| 24.2 | 0.0500 | ||
| 25.2 | 0.0300 | ||
| 26.2 | 0.0333 | ||
| 27.2 | 0.0004 | ||
| 25 | VGL0910 | 13 | 0.0038 |
| 14 | 0.0096 | ||
| 16.1 | 0.0921 | ||
| 17 | 0.0063 | ||
| 17.1 | 0.1008 | ||
| 18 | 0.0246 | ||
| 18.1 | 0.0617 | ||
| 19.1 | 0.1700 | ||
| 20.1 | 0.0404 | ||
| 20.3 | 0.0050 | ||
| 21.1 | 0.3154 | ||
| 22 | 0.0325 | ||
| 22.1 | 0.1058 | ||
| 23 | 0.0154 | ||
| 23.1 | 0.0167 | ||
| 26 | VGL1063 | 8 | 0.0233 |
| 12 | 0.0842 | ||
| 13 | 0.0396 | ||
| 14 | 0.1983 | ||
| 15 | 0.1029 | ||
| 16 | 0.0071 | ||
| 17 | 0.0875 | ||
| 18 | 0.0688 | ||
| 19 | 0.0438 | ||
| 20 | 0.0863 | ||
| 21 | 0.0892 | ||
| 22 | 0.1567 | ||
| 23 | 0.0125 | ||
| 27 | VGL1165 | 14 | 0.0008 |
| 15 | 0.0004 | ||
| 16 | 0.0058 | ||
| 17 | 0.0292 | ||
| 18 | 0.1113 | ||
| 19 | 0.0154 | ||
| 20 | 0.0292 | ||
| 21 | 0.1121 | ||
| 22 | 0.0004 | ||
| 23 | 0.0246 | ||
| 25 | 0.0038 | ||
| 26 | 0.3533 | ||
| 27 | 0.0708 | ||
| 28 | 0.1733 | ||
| 29 | 0.0079 | ||
| 30 | 0.0121 | ||
| 31 | 0.0233 | ||
| 32 | 0.0104 | ||
| 33 | 0.0150 | ||
| 34 | 0.0008 | ||
| 28 | VGL1828 | 15 | 0.0146 |
| 16 | 0.3088 | ||
| 17 | 0.0050 | ||
| 18 | 0.0213 | ||
| 19 | 0.0325 | ||
| 19.3 | 0.0029 | ||
| 20 | 0.1088 | ||
| 20.3 | 0.0004 | ||
| 21 | 0.3600 | ||
| 22 | 0.0633 | ||
| 23 | 0.0817 | ||
| 24 | 0.0008 | ||
| 29 | VGL2009 | 9 | 0.0021 |
| 10 | 0.0025 | ||
| 11 | 0.1454 | ||
| 12 | 0.0254 | ||
| 13 | 0.2138 | ||
| 14 | 0.2283 | ||
| 15 | 0.2958 | ||
| 16 | 0.0854 | ||
| 17 | 0.0013 | ||
| 30 | VGL2409 | 14 | 0.0117 |
| 15 | 0.0700 | ||
| 16 | 0.3267 | ||
| 17 | 0.3254 | ||
| 18 | 0.2125 | ||
| 19 | 0.0029 | ||
| 20 | 0.0417 | ||
| 21 | 0.0092 | ||
| 31 | VGL2918 | 12 | 0.0446 |
| 13 | 0.2804 | ||
| 14 | 0.1496 | ||
| 15 | 0.0683 | ||
| 16 | 0.0108 | ||
| 16.3 | 0.0054 | ||
| 17.3 | 0.0108 | ||
| 18.3 | 0.0667 | ||
| 19.3 | 0.0408 | ||
| 20.3 | 0.0304 | ||
| 21.3 | 0.0667 | ||
| 22.3 | 0.2171 | ||
| 23.3 | 0.0083 | ||
| 32 | VGL3008 | 10 | 0.0004 |
| 13 | 0.1283 | ||
| 14 | 0.0083 | ||
| 15 | 0.0704 | ||
| 16 | 0.0263 | ||
| 17 | 0.1104 | ||
| 18 | 0.1542 | ||
| 19 | 0.1771 | ||
| 20 | 0.2033 | ||
| 21 | 0.1038 | ||
| 22 | 0.0104 | ||
| 23 | 0.0071 | ||
| 33 | VGL3235 | 11 | 0.0113 |
| 12 | 0.0188 | ||
| 13 | 0.1263 | ||
| 14 | 0.2783 | ||
| 15 | 0.3433 | ||
| 16 | 0.0288 | ||
| 17 | 0.0396 | ||
| 18 | 0.0658 | ||
| 19 | 0.0833 | ||
| 20 | 0.0046 |
Standard genetic assessment based on 33 autosomal STR loci
| N | Na | Ne | Ho | He | F | ||
|---|---|---|---|---|---|---|---|
| Mean | 1200 | 9.515 | 4.477 | 0.731 | 0.748 | 0.023 | |
| SE | 0.535 | 0.268 | 0.016 | 0.016 | 0.004 |
Standard genetic assessment based on 7 STRs in the DLA region
| N | Na | Ne | Ho | He | F | ||
|---|---|---|---|---|---|---|---|
| Mean | 1200 | 8.857 | 3.840 | 0.670 | 0.690 | 0.030 | |
| SE | 0.999 | 0.600 | 0.047 | 0.047 | 0.005 |
Standard genetic assessment for individual STR loci
Havanese
| # | Locus | N | Na | Ne | Ho | He | F |
|---|---|---|---|---|---|---|---|
| 1 | AHT121 | 1200 | 12 | 3.132 | 0.676 | 0.681 | 0.007 |
| 2 | AHT137 | 1200 | 11 | 6.432 | 0.833 | 0.845 | 0.014 |
| 3 | AHTH130 | 1200 | 9 | 4.051 | 0.716 | 0.753 | 0.050 |
| 4 | AHTh171-A | 1200 | 10 | 4.849 | 0.786 | 0.794 | 0.010 |
| 5 | AHTh260 | 1200 | 9 | 4.840 | 0.770 | 0.793 | 0.029 |
| 6 | AHTk211 | 1200 | 5 | 3.145 | 0.653 | 0.682 | 0.043 |
| 7 | AHTk253 | 1200 | 5 | 2.113 | 0.485 | 0.527 | 0.079 |
| 8 | C22.279 | 1200 | 8 | 4.633 | 0.758 | 0.784 | 0.034 |
| 9 | FH2001 | 1200 | 9 | 2.634 | 0.626 | 0.620 | -0.009 |
| 10 | FH2054 | 1200 | 10 | 6.194 | 0.833 | 0.839 | 0.007 |
| 11 | FH2848 | 1200 | 9 | 4.400 | 0.753 | 0.773 | 0.026 |
| 12 | INRA21 | 1200 | 7 | 4.418 | 0.743 | 0.774 | 0.039 |
| 13 | INU005 | 1200 | 9 | 2.463 | 0.595 | 0.594 | -0.002 |
| 14 | INU030 | 1200 | 7 | 2.249 | 0.545 | 0.555 | 0.019 |
| 15 | INU055 | 1200 | 6 | 4.110 | 0.730 | 0.757 | 0.035 |
| 16 | LEI004 | 1200 | 9 | 2.111 | 0.535 | 0.526 | -0.017 |
| 17 | REN105L03 | 1200 | 9 | 3.464 | 0.685 | 0.711 | 0.037 |
| 18 | REN162C04 | 1200 | 6 | 3.917 | 0.731 | 0.745 | 0.019 |
| 19 | REN169D01 | 1200 | 8 | 3.930 | 0.734 | 0.746 | 0.015 |
| 20 | REN169O18 | 1200 | 7 | 5.201 | 0.793 | 0.808 | 0.018 |
| 21 | REN247M23 | 1200 | 7 | 3.547 | 0.703 | 0.718 | 0.021 |
| 22 | REN54P11 | 1200 | 7 | 5.152 | 0.773 | 0.806 | 0.041 |
| 23 | REN64E19 | 1200 | 9 | 3.489 | 0.712 | 0.713 | 0.002 |
| 24 | VGL0760 | 1200 | 14 | 7.196 | 0.852 | 0.861 | 0.011 |
| 25 | VGL0910 | 1200 | 15 | 6.023 | 0.813 | 0.834 | 0.025 |
| 26 | VGL1063 | 1200 | 13 | 8.806 | 0.877 | 0.886 | 0.011 |
| 27 | VGL1165 | 1200 | 20 | 5.304 | 0.763 | 0.811 | 0.060 |
| 28 | VGL1828 | 1200 | 12 | 4.013 | 0.763 | 0.751 | -0.017 |
| 29 | VGL2009 | 1200 | 9 | 4.663 | 0.758 | 0.786 | 0.036 |
| 30 | VGL2409 | 1200 | 8 | 3.779 | 0.713 | 0.735 | 0.030 |
| 31 | VGL2918 | 1200 | 13 | 6.002 | 0.826 | 0.833 | 0.009 |
| 32 | VGL3008 | 1200 | 12 | 7.054 | 0.849 | 0.858 | 0.011 |
| 33 | VGL3235 | 1200 | 10 | 4.435 | 0.733 | 0.775 | 0.053 |
Standard genetic assessment for 7 STRs in the DLA region
Havanese
| # | Locus | N | Na | Ne | Ho | He | F |
|---|---|---|---|---|---|---|---|
| 1 | DLA I-3CCA | 1200 | 12 | 5.046 | 0.783 | 0.802 | 0.023 |
| 2 | DLA I-4ACA | 1200 | 12 | 4.703 | 0.761 | 0.787 | 0.034 |
| 3 | DLA I-4BCT | 1200 | 8 | 2.044 | 0.503 | 0.511 | 0.016 |
| 4 | DLA1131 | 1200 | 11 | 6.660 | 0.833 | 0.850 | 0.019 |
| 5 | 5ACA | 1200 | 6 | 3.590 | 0.701 | 0.721 | 0.029 |
| 6 | 5ACT | 1200 | 5 | 2.130 | 0.501 | 0.530 | 0.056 |
| 7 | 5BCA | 1200 | 8 | 2.708 | 0.608 | 0.631 | 0.036 |